Biotopes typical of intertidal mud and sandflats.

Biotope definition and code Environmental Characteristics Characterising species
Hediste (Nereis) diversicolor and Macoma balthica in sandy mud shores

(LMU.HedMac)

 

Subtypes of this biotope are:

LMU.HedMac.Are - least sheltered, abundant Arenicola marina and frequent Cerastoderma edule;

LMU.HedMac.Pyg - contains less A. marina; LMU.HedMac.Mare - contains Mya arenaria in high densities

Full-Variable salinity

Sheltered, Very sheltered, Extremely sheltered

Sandy mud-Mud

 

Eulittoral

Mid shore, Lower shore

Anoxic layer present

• Polychaetes, typically Hediste (Nereis) diversicolor

• Other smaller polychaetes include Eteone longa and Nephtys hombergii, Tharyx marioni, Pygospio elegans, Arenicola marina and Manayunkia aestuarina.

•Oligochaete worms include Tubificoides spp.

•Amphipod, Corophium volutator

•Mud snail, Hydrobia ulvae

•Bivalves include Macoma balthica

•Green algae, e.g. Enteromorpha

Hediste diversicolor and Scrobicularia plana in reduced salinity mud shores

(LMU.HedScr)

 

 

Variable-Reduced/Low salinity

Sheltered, Very sheltered, Extremely sheltered

Mud-Sandy mud

Eulittoral

Upper shore, Mid shore, Lower shore

Anoxic layer present

•Hediste (Nereis) diversicolor and the bivalve Scrobicularia plana are abundant

•Other polychaetes include Eteone longa

•Oligochaete, Tubificoides benedeni

•Isopod, Cyathura carinata

•Other bivalves include Macoma balthica and Cerastoderma edule

Hediste (Nereis) diversicolor and Streblospio shrubsolii in sandy mud or soft mud shores

(LMU.HedStr)

 

 

Variable-Low salinity

Very sheltered-Extremely sheltered

Mud-Sandy mud

Eulittoral

Mid shore, Lower shore

Black, possibly nutrient enriched

Streblospio shrubsolii, Tharyx killariensis and Manayunkia aestuarina

Hediste (Nereis) diversicolor, Nephtys hombergii, Pygospio elegans

Corophium volutator, Hydrobia ulvae, Macoma balthica and Abra tenuis

Tubificoides spp.

Hediste (Nereis) diversicolor and oligochaetes in low salinity mud shores

(LMU.HedOl)

 

 

 

Reduced-Low salinity

Extremely sheltered

Mud-Sandy mud

Littoral fringe, Eulittoral

Upper shore, Mid shore, Lower shore

•Oligochaetes, including Tubificoides spp. and Hediste (Nereis) diversicolor are abundant

Corophium volutator

•Reduced polychaetes and absence of bivalves

Barren coarse sands

(LGS.BarSnd)

Full salinity

Exposed-mod. Exposed

Coarse-medium sand

Supralittoral, Eulittoral

Strandline, Upper shore, Mid shore, Lower shore

•Sparse macrofauna

•Low abundances of burrowing amphipods Bathyporeia spp. or Pontocrates spp. and Eurydice pulchra

Burrowing amphipods and Eurydice pulchra in well drained clean sand shores

(LGS.AEur)

Full salinity

Exposed-Moderately exposed

Medium sand

Eulittoral

Upper shore, Mid shore, Lower shore

•Burrowing amphipods

Eurydice pulchra

•Impoverished polychaetes only Scolelepis squamata

Burrowing amphipods and polychaetes in clean sand shores

(LGS.AP)

 

 

Sub biotopes are (LGS.AP.P.) and (LGS.AP.Pon), depending on the amphipod to polychaete ratio

Full salinity

Exposed, Mod. Exposed, Sheltered

Medium-Fine sand

Eulittoral

Mid shore, Lower shore

•Burrowing amphipods and polychaetes includes Pontocrates and Bathyporeia spp. and Nephtys cirrosa, Scolelepis squamata, Paraonis fulgens and Arenicola marina.

•Occasional bivalves e.g. Angulus tenuis

•Isopod, Eurydice pulchra

Dense Lanice conchilega in tide-swept lower shore sand

(LGS.Lan)

Full-variable salinity

Moderately exposed Sheltered, Very sheltered

Strong-Moderately strong tidal streams

Medium sand-fine sand

Eulittoral

Mid shore, Lower shore

•Dense populations of Lanice conchilega

•Other polychaetes include Nephtys cirrus, Nephtys hombergii and Pygospio elegans

•Few crustaceans

•Bivalve cockles, Cerastoderma edule

Bathyporeia pilosa and Corophium spp. in upper shore slightly muddy fine sand shores

(LMS.BatCor)

 

 

Variable salinity

Mod. exposed, Sheltered, Very sheltered

Muddy fine sand

Eulittoral

Upper shore, Mid shore

•Amphipods, Bathyporeia pilosa, Corophium arenarium and Corophium volutator

•Polychaetes and bivalves (with the exception of Macoma balthica) are limited in their abundance and variety.

Polychaetes and Cerastoderma edule in fine sand or muddy and shores

(LMS.PCer)

Full salinity

Moderately exposed-Sheltered

Fine sand or muddy sand

Eulittoral

Mid shore, Lower shore

Cerastoderma edule and other bivalves

•Reduced amphipod populations, Bathyporeia sarsi

•Polychaetes, Nephtys hombergii, Scoloplos armiger, Pygospio elegans, Spio filicornis and Capitella capitata

•Oligochaetes

Macoma balthica and Arenicola marina in muddy sand shores

(LMS.MacAre)

 

 

Full-Variable salinity

Mod.exposed, Sheltered, Very sheltered, Extremely sheltered

Fine sand or muddy sand

Eulittoral

Upper shore, Mid shore, Lower shore

Anoxic layer present

Arenicola marina and Scoloplos armiger

Macoma balthica and Cerastoderma edule

Corophium volutator can be common

References