Key references for background reading on the ecology of brittlestar beds are shown in boldface.

Allain, J.-Y., 1974. Ecologie des bancs d’Ophiothrix fragilis (Abildgaard) (Echinodermata, Ophiuroidea) dans le golfe normanno-breton. Cahiers de Biologie Marine, 15: 255-273.

Allain, J.-Y., Romano, J.C. & Semroud, R., 1978. Ophiocomina nigra (O.F. Mueller) (Echinodermata, Ophiuroidea) dans la region d’Alger. Pelagos, 5: 50-64.

Allen, E.J., 1899. On the fauna and bottom-deposits near the thirty-fathom line from the Eddystone Grounds to Start Point. Journal of the Marine Biological Association of the United Kingdom, 5: 365-542.

Aronson, R.B., 1987a. Predation on fossil and recent ophiuroids. Paleobiology, 13: 187-192.

Aronson, R.B., 1987b. A murder mystery from the Mesozoic. New Scientist, 1581: 56-59.

Aronson, R.B. & Sues, H.-D., 1988. The fossil record of brittlestar beds. In: Echinoderm Biology. Proceedings of the 6th International Echinoderm Conference, Victoria, August 1987. eds. Burke, R.D., Mladenov, P.V., Lambert, P. & Parsley, R.L. Rotterdam, A.A. Balkema, pp. 147-148.

Auster, P.J., 1993. ROVs for science. Marine Technology Society Journal, 26: 3-4.

Black, K.D. (ed.) 1996. Aquaculture and sea lochs. Scottish Association for Marine Science, Oban.

Broom, D.M., 1975. Aggregation behaviour of the brittle-star Ophiothrix fragilis. Journal of the Marine Biological Association of the United Kingdom, 55: 191-197.

Brun, E., 1972. Food and feeding habits of Luidia ciliaris (Echinodermata: Asteroidea). Journal of the Marine Biological Association of the United Kingdom, 52: 225-236.

Bunker, F. St P.D. & Hiscock, K.S., 1987. Sublittoral habitats, communities and species of the Skomer Marine Reserve - a review (Contractor, Field Studies Council, Pembroke). Nature Conservancy Council CSD Report No. 747 (FSC Report No. (OFC)/1/87).

Carlton, J.T., 1996. Marine bioinvasions: the alteration of marine ecosystems by nonindigenous species. Oceanography, 9: 36-42.

Chadwick, H.C., 1886. Report on the Ophiuroidea of the LMBC district. In: Herdman, W.A. (ed.) First Report on the fauna of Liverpool Bay and neighbouring seas. pp. 140- 143.

Connor, D.W., 1989. Survey of Loch Duich, Loch Long and Loch Alsh. Marine Nature Conservation Review Report MNCR/SR/010/89 (Nature Conservancy Council CSD Report No. 977).

Connor, D.W., Dalkin, M.J., Hill, T.O., Holt, R.H.F. & Sanderson, W.G., 1997. Marine Nature Conservation Review: Marine biotope classification for Britain and Ireland. Volume 2. Sublittoral biotopes. Version 97.06. Joint Nature Conservation Committee Report No. 230.

Covey, R., 1991. Benthic marine ecosystems in Great Britain: a review of current knowledge. Eastern England and English Channel (MNCR Coastal Sectors 6 and 7). Nature Conservancy Council CSD Report No. 1172 (MNCR Report No. MNCR/OR/08).

Covey, R., 1992. Sublittoral survey of the north coast of the outer Solway (Mull of Galloway to Auchencairn). Nature Conservancy Council CSD Report No. 1193 (MNCR Report MNCR/SR/15).

Cutts, N. & Hemingway, K., 1996. The Solway Firth: broad scale habitat mapping. Scottish Natural Heritage Research, Survey and Monitoring Report No. 46.

Czihak, G., 1954. Die Schlangensterne einer submersen Höhle des Tyrrhenischen Meeres. Österreicher Zoologische Zeitschrifft, 5: 15-29.

Dahm, C., 1993. Growth, production and ecological significance of Ophiura albida and O. ophiura (Echinodermata: Ophiuroidea) in the German Bight. Marine Biology, 116: 431-437.

Davies, J. & Hall-Spencer, J., 1996. Mapping of the benthic biotopes in the proposed Sound of Arisaig Special Area of Conservation. Scottish Natural Heritage Research, Survey and Monitoring Report No. 83.

Davoult, D. 1990. Biofacies et structure trophique du peuplement des cailloutis du Pas-de- Calais (France). Oceanologia Acta, 13: 335-348.

Davoult, D. & Gounin, F., 1995b. Nitrogen excretion by a dense population of Ophiothrix fragilis (Abildgaard): role in the exchanges at the water-sediment interface. In: Echinoderm Research 1995. Proceedings of the 4th European Echinoderms Colloquium, London, 10-13 April 1995. eds. Emson, R., Smith, A. & Campbell, A. Rotterdam: A.A. Balkema, pp. 65-69.

Davoult, D., Gounin, F. & Janquin, M.K., 1991. Ammonium excretion by the ophiuroid Ophiothrix fragilis as a function of season and tide. Marine Biology, 111: 425-429.

Davoult, D., Gounin, F. & Richard, A., 1990. Dynamique et reproduction de la population d’Ophiothrix fragilis (Abildgaard) du detroit du Pas de Calais (Manche Orientale). Journal of Experimental Marine Biology and Ecology, 138: 201-216.

Devon Wildlife Trust, 1993. Marine survey report. Plymouth Sound and approaches. A report on the wildlife resource of Plymouth Sound and the adjacent open coast.

Dipper, F.A., Irving, R.A. & Fowler, S.L., 1989. Sublittoral survey of the Wash by diving and dredging (1985 and 1986). Nature Conservancy Council CSD Report No. 976.

Downie, A.J., 1990. The reproduction and development of flatfish in the proximity of the sludge dump site in the Firth of Clyde. Ph.D Thesis, University of London.

Dyrynda, P., 1984. Investigation of the subtidal ecology of the Fleet Lagoon (Dorset). Nature Conservancy Council CSD Report No. 530.

Eno, N.C., Clark, R.A. & Sanderson, W.G. (eds.), 1997. Non-native species in British waters: a review and directory. Joint Nature Conservation Committee, Peterborough.

Entec, 1996. Broad scale habitat mapping of intertidal and subtidal coastal areas: Loch Maddy, North Uist. Scottish Natural Heritage Research, Survey and Monitoring Report No. 76.

Erwin, D.G., 1977. A diving survey of Strangford Lough: the benthic communities and their relation to substrate - a preliminary account. In: Biology of Benthic Organisms. Proceedings 11th European Marine Biology Symposium, Galway, 1976. eds. Keegan, B.F., Ó Céidigh, P., & Boaden, P.J.S., Oxford, Pergamon Press, pp. 215-224.

Feder, H.M., 1981. Aspects of the feeding biology of the brittlestar Ophiura texturata. Ophelia, 20: 215-235.

Fedra, K., Ölscher, E.M., Scherübel, C., Stachowitsch, M. & Wurzian, R.S., 1976. On the ecology of a North Adriatic benthic community: distribution, standing crop and composition of the macrobenthos. Marine Biology, 38: 129-145.

Fontaine, A.R., 1962. The colours of Ophiocomina nigra (Abildgaard) I. Colour variation and its relation to distribution. Journal of the Marine Biological Association of the United Kingdom, 42: 1-8.

Fontaine, A.R., 1964. The integumentary mucous secretions of the ophiuroid Ophiocomina nigra. Journal of the Marine Biological Association of the United Kingdom, 44: 145-162.

Fontaine, A.R., 1965. The feeding mechanism of the ophiuroid Ophiocomina nigra. Journal of the Marine Biological Association of the United Kingdom, 45: 373-385.

Foster-Smith, R.L., Sotheran, I., Foster-Smith, J.L. & Bunker, F., 1996. Mapping survey of the sublittoral and littoral biotopes of the Berwickshire coast. Scottish Natural Heritage Research, Survey and Monitoring Report, No. 60.

Foster-Smith, R.L., Sotheran. I. & Walton, R., 1997. Broadscale mapping of the habitats and biota of the sublittoral seabed of the Wash. Report to English Nature.

Fratt, D.B. & Dearborn, J.H., 1984. Feeding biology of the Antarctic brittle star Ophionotus victoriae (Echinodermata: Ophiuroidea). Polar Biology, 3: 127-139.

Fujita, T. & Ohta, S., 1990. Size structure of dense populations of the brittle star Ophiura sarsii (Ophiuroidea: Echinodermata) in the bathyal zone around Japan. Marine Ecology Progress Series, 64: 113-122.

Gage, J.D., 1990. Skeletal growth bands in brittlestars: microstructure and significance as age markers. Journal of the Marine Biological Association of the United Kingdom, 70: 209-224.

Gage, J.D. & Tyler, P.A., 1991. Deep-sea biology: a natural history of organisms at the deep-sea floor. Cambridge University Press.

Garner, R., 1878. Professor Edward Forbes and his country. Midland Naturalist, 1: 67-70.

George, C.L. & Warwick, R.M., 1985. Annual macrofauna production in a hard-bottom reef community. Journal of the Marine Biological Association of the United Kingdom, 65: 713-736.

Gorzula, S.J., 1976. The distribution of epi-benthic ophiuroids in Cumbrae waters. The Western Naturalist, 5: 71-80.

Gorzula, S.J., 1977. A study of growth in the brittle-star Ophiocomina nigra. The Western Naturalist, 6: 13-33.

Gounin, F., Davoult, D. & Richard, A., 1995. Role of a dense bed of Ophiothrix fragilis (Abildgaard) in the transfer of heavy metals at the water-sediment interface. Marine Pollution Bulletin, 30: 736-741.

Guille, A., 1964. Contribution a l’étude de la systématique et de l’écologie d’Ophiothrix quinquemaculata (D. Ch.). Vie et Milieu, 15: 243-308.

Hagen, N.T., 1995. Sea urchin outbreaks and epizootic disease as regulating mechanisms in coastal ecosystems. In: Biology and Ecology of Shallow Coastal Waters. Proceedings 28th European Marine Biology Symposium, Iraklio, Crete, 1993. eds. Eleftheriou, A., Ansell, A.D. & Smith, C.J., Fredensborg, Olsen & Olsen, pp. 303-308.

Hiscock, K., 1984. Sublittoral survey of Bardsey and the Lleyn Peninsula, August 13th to 27th 1983. Volume 1, Survey Report. Report to the Nature Conservancy Council from the Field Studies Council Oil Pollution Research Unit, Orielton, Pembroke, Dyfed.

Hiscock, K., submitted. Identifying marine ‘sensitive areas’ - the importance of understanding life-cycles. Paper submitted for publication in the proceedings of the symposium on Aquatic life cycle strategies: survival in a variable environment, April 1997.

Hiscock, K. & Moore, J., 1986. Surveys of harbours, rias and estuaries in southern Britain: Plymouth area including the Yealm. Volume 1, Report. Report to the Nature Conservancy Council from the Field Studies Council Oil Pollution Research Unit, Orielton, Pembroke, Dyfed.

Holme, N.A., 1971. Macrofauna sampling. In: IBP Handbook No. 16, Methods for the study of marine benthos, 2nd edition. eds. Holme, N.A. & McIntyre, A.D.. Blackwell Scientific Publications, Oxford & Edinburgh. pp. 80-130.

Howson, C.M., 1988. Marine Nature Conservation Review. Survey of Shetland, Foula and Fair Isle, 1987. Volume 1, Report. Report to the Nature Conservancy Council from the Field Studies Council Oil Pollution Research Unit, Orielton, Pembroke, Dyfed.

Howson, C.M., 1990. Marine Nature Conservation Review Surveys of Scottish sealochs. Sealochs of Arisaig and Moidart. Report to the Joint Nature Conservation Committee from the University Marine Biological Station, Millport.

Howson, C.M., Connor, D.W. & Holt, R.H.F., 1994. The Scottish sealochs - an account of surveys undertaken for the Marine Nature Conservation Review. Joint Nature Conservation Committee Report No. 164 (Marine Nature Conservation Review Report MNCR/SR/27).

Hutchins, D.A., Boisson, F., Fowler, S.W. & Fisher, N.S., 1996. Temperature effects on uptake and retention of contaminant radionuclides and trace metals by the brittlestar Ophiothrix fragilis. Marine Environmental Research, 41: 363-378.

Keegan, B.F., O’Connor, B.D.S. & Könnecker, G., 1985. Littoral and benthic investigations on the west coast of Ireland - XX. Echinoderm aggregations. Proceedings of the Royal Irish Academy, 85B: 91-99.

Keegan, B.F. & Mercer, J.P., 1986. An oceanographical survey of Killary Harbour on the west coast of Ireland. Proceedings of the Royal Irish Academy, 86B: 1-70.

Kelly, M.S. & McKenzie, J.D., 1995. A survey of the occurrence and morphology of sub-cuticular bacteria in shelf echinoderms from the north-east Atlantic. Marine Biology, 123: 741-756.

Koehler, R., 1921: Echinodermes. Faune de France, 1. Paris: Office Centrale de Faunistique.

Leewis, R.J., Waardenburg, H.W. & van der Tol, M.W.M., 1994. Biomass and standing stock on sublittoral hard substrates in the Oosterschelde Estuary (SW Netherlands). Hydrobiologia, 282/283: 397-412.

Lessios, H.A., 1988. Mass mortality of Diadema antillarum in the Caribbean: what have we learned? Annual Review of Ecology and Systematics, 19: 371-393.

Lindley, J.A., Gamble, J.C. & Hunt, H.G., 1995. A change in the zooplankton of the central North Sea (55^o to 58^o N): a possible consequence of changes in the benthos. Marine Ecology Progress Series, 119: 299-303.

Mackie, A.S.Y., Oliver, P.G. & Rees, E.I.S., 1995. Benthic biodiversity in the southern Irish Sea. Studies in marine biodiversity and systematics from the National Museum of Wales. BIOMÔR Reports.

Magorrian, B.H., Service, M. & Clarke, W., 1995. An acoustic bottom classification of Strangford Lough, Northern Ireland. Journal of the Marine Biological Association of the United Kingdom, 75: 987-982.

Migné, A. & Davoult, D., 1997. Distribution quantitative de la macrofaune benthique du peuplement des cailloutis dans le detroit du Pas de Calais (Manche Orientale, France). Oceanologia Acta, 20: 453-460.

Mills, J.L., 1997.Chapter 11. Liverpool Bay and the Solway (Rhôs-on-Sea to the Mull of Galloway) (MNCR Sector 11). In: Marine Nature Conservation Review. Benthic marine ecosystems: a review of current knowledge for Great Britain and the north-east Atlantic. ed. Hiscock, K., Peterborough, Joint Nature Conservation Committee (Coasts and Seas of the United Kingdom. MNCR Series).

Morison, G.W., 1979. Studies on the ecology of the sub-Antarctic ophiuroid Ophionotus hexactis (E.A. Smith). Ph.D Thesis, University of London.

Newton, L.C., 1995. The effects of natural and induced stress on echinoderms. Ph.D Thesis, Napier University, Edinburgh.

Newton, L.C. & McKenzie, J.D., 1995. Echinoderms and oil pollution: a potential stress assay using bacterial symbionts. Marine Pollution Bulletin, 31: 453-456.

Ott, J. & Fedra, K., 1977. Stabilising properties of a high-biomass benthic community in a fluctuating ecosystem. Helgolander Wissenschaftliche Meeresuntersuchungen 30: 485-494.

Pagett, R., 1984. The sublittoral ecology of the St Abbs area, Berwickshire (III). Report to the Nature Conservancy Council No. 512, from the Marine Conservation Society.

Picton, B.E., 1993. A field guide to the shallow-water echinoderms of the British Isles. London, Immel Publishing Ltd.

Raymont, J.E.G., 1950. A fish cultivation experiment in an arm of a sea loch IV. The bottom fauna of Kyle Scotnish. Proceedings of the Royal Society of Edinburgh (B), 64: 65- 108.

Reese, E.S., 1966. The complex behaviour of echinoderms. In: Physiology of Echinodermata ed. Boolootian, R.A., New York, John Wiley & Sons, pp. 157-218.

Roushdy, H.M. & Hansen, V.K., 1960. Ophiuroids feeding on phytoplankton. Nature, 188: 517-518.

Russell, F.S., 1935. On the value of certain plankton animals as indicators of water movements in the English Channel and North Sea. Journal of the Marine Biological Association of the United Kingdom, 20: 309-332.

Saldanha, L., 1973. Présence de populations dense d’Ophiocomina nigra (O.F. Müller) sur la côte d’Arrábida (Portugal). Revista da Faculdade de Ciencias de Lisboa, Série 2C, 17: 427-436.

Service, M., 1993. The structure and chemistry of the superficial sediments of Strangford Lough, Northern Ireland. Marine Pollution Bulletin, 26: 343-345.

Service, M. & Magorrian, B.H., 1997. The extent and temporal variation of disturbance to epibenthic communities in Strangford Lough, Northern Ireland. Journal of the Marine Biological Association of the United Kingdom, 77: 1151-1164.

Smith, J.E., 1938. Occurrence of young Ophiothrix fragilis within the genital bursae of the adults. Nature, 141: 544.

Smith, J.E., 1940. The reproductive system and associated organs of the brittle-star Ophiothrix fragilis. Quarterly Journal of Microscopic Science, 132: 267-309.

Stachowitsch, M., 1984. Mass mortality in the Gulf of Trieste: the course of community destruction. P.S.Z.N.I. Marine Ecology, 5: 243-264.

Southward, A.J., 1980. The western English Channel - an inconstant ecosystem? Nature, London, 285: 361-366.

Tyler, P.A., 1976. The ecology and reproductive biology of the genus Ophiura, with special reference to the Bristol Channel. Ph.D Thesis, University of Wales, Swansea.

Tyler, P.A., 1977. Seasonal variation and ecology of gametogenesis in the genus Ophiura (Ophiuroidea: Echinodermata) from the Bristol Channel. Journal of Experimental Marine Biology and Ecology, 30: 185-197.

Vermeij, G.J., 1977. The Mesozoic marine revolution: evidence from snails, predators and grazers. Paleobiology, 3: 245-258.

Vermeij, G.J., 1987. Evolution and escalation: an ecological history of life. Princeton University Press, Princeton, New Jersey.

Vevers, H.G., 1952. A photographic survey of certain areas of the sea floor near Plymouth. Journal of the Marine Biological Association of the United Kingdom, 31: 215-222.

Wilding, T.A. & Gage, J.D., 1995. Skeletal growth marks in the brittle star Ophiura ophiura (Linnaeus): do they reflect a seasonal growth pattern? In: Echinoderm Research 1995. Proceedings of the 4th European Echinoderms Colloquium, London, 10-13 April 1995. eds. Emson, R., Smith, A. & Campbell, A. Rotterdam: A.A. Balkema, pp. 283-291.

Warner, G.F. & Woodley, J.D., 1975. Suspension-feeding in the brittle-star Ophiothrix fragilis. Journal of the Marine Biological Association of the United Kingdom, 55: 199-210.

Wolff, W.J., 1968. The Echinodermata of the estuarine region of the rivers Rhine, Meuse and Scheldt, with a list of species occurring in the coastal waters of the Netherlands. Netherlands Journal of Sea Research, 4: 59-85.